In this audio slide show, meet Kanzi, the Trust's alpha male, who has demonstrated an extraordinary ability to communicate on our terms.—Rima Chaddha ...

Kanzi has died at the age of 44. The iconic bonobo reshaped our understanding of ape intelligence, challenging long-held beliefs about what separates humans from our closest living relatives.

Kanzi was not the first great ape to learn how to communicate with humans using symbols. Koko the gorilla and Washoe the chimpanzee learned signs that were adapted from American Sign Language. But ...